Ruff
Calidris pugnax
- KingdomAnimalia (Animals)
- PhylumChordata (Chordates)
- ClassAves (Birds)
- OrderCharadriiformes (Shorebirds and Allies)
- FamilyScolopacidae (Sandpipers and Allies)
- GenusCalidris (Calidris Sandpipers)
- SpeciesCalidris pugnax (Ruff)
About the Ruff
The ruff (Calidris pugnax) is a medium-sized wading bird that breeds in marshes and wet meadows across northern Eurasia. This highly gregarious sandpiper is migratory and sometimes forms huge flocks in its winter grounds, which include southern and western Europe, Africa, southern Asia and Australia.
The ruff is a long-necked, pot-bellied bird. This species shows marked sexual dimorphism; the male is much larger than the female (the reeve), and has a breeding plumage that includes brightly coloured head tufts, bare orange facial skin, extensive black on the breast, and the large collar of ornamental feathers that inspired this bird's English name. The female and the non-breeding male have grey-brown upperparts and mainly white underparts. Three differently plumaged types of male, including a rare form that mimics the female, use a variety of strategies to obtain mating opportunities at a lek, and the colourful head and neck feathers are erected as part of the elaborate main courting display. The female has one brood per year and lays four eggs in a well-hidden ground nest, incubating the eggs and rearing the chicks, which are mobile soon after hatching, on her own. Predators of wader chicks and eggs include mammals such as foxes, feral cats and stoats, and birds such as large gulls, crows and skuas.
The ruff forages in wet grassland and soft mud, probing or searching by sight for edible items. It primarily feeds on insects, especially in the breeding season, but it will consume plant material, including rice and maize, on migration and in winter. Classified as "least concern" on the IUCN Red List criteria, the global conservation concerns are relatively low because of the large numbers that breed in Scandinavia and the Arctic. However, the range in much of Europe is contracting because of land drainage, increased fertiliser use, the loss of mown or grazed breeding sites, and over-hunting. This decline has seen it listed in the Agreement on the Conservation of African-Eurasian Migratory Waterbirds (AEWA). Wikipedia →
Closest Relatives at Manas National Park
Temminck's Stint
Calidris temminckii
Green Sandpiper
Tringa ochropus
Common Sandpiper
Actitis hypoleucos
Common Snipe
Gallinago gallinago
Common Greenshank
Tringa nebularia
Common Redshank
Tringa totanus
Pin-tailed Snipe
Gallinago stenura
Wood Sandpiper
Tringa glareola
Marsh Sandpiper
Tringa stagnatilis
Spotted Redshank
Tringa erythropus
Red-wattled Lapwing
Vanellus indicus
River Lapwing
Vanellus duvaucelii
Global Distribution
Where this species is recorded worldwide. Restricted to human observations from 2019 onwards — the same kind of record and the same period the probabilities on this page are built from, so collar-tracking studies, camera-trap arrays and museum specimens are excluded. Around Santa Rosa that removes 5,924 GPS fixes from a handful of tagged jaguars.
where this species is recorded (GBIF density — it tracks where observers go, not how likely a sighting is) · parks where it occurs — tap a pin for the sighting probability there